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  • TITLE
  • CERTIFICATE
  • DECLARATION
  • ACKNOWLEDGEMENT
  • CONTENTS
  • I. Introduction
  • PLATE - I TROPIDONOTUS, Natrix uiscator piscator Schneider.
  • PLATE - II RATSNAKE, Ptyas mucasus Linne
  • PLATE - III COBRA, Naja naja naja L.inne
  • Tropidonotus
  • Ratsnake
  • Cobra
  • II. Materials and methods
  • Food -getting and feeding behaviour
  • Epidermal structure of the tongue
  • Osteological elements
  • Quantitative analysis of the bony elements
  • Jaw-muscles
  • A. Disposition and preservation
  • B. Quantitative myology
  • C. The force of adduction
  • D. The force of abduction
  • PLATE - IV Diagrammatic representation of the lever systems involving the opening and closing of the jaws.
  • E. The force of protraction
  • F. The force of retraction
  • Histochemical observations
  • III. Food-getting and feeding behaviour
  • Field observations
  • Analysis of stomach contents
  • IV. Epidermal structure of the tongue
  • PLATE - V Fig.1, 2 & 3 Diagrammatic representation of snakes tounge. Fig. 4. 5 & 6 Scanning electron micrographs showing the epidermal structure of the tongue of tropjdnnotus.
  • PLATE - VI Scanning electron micrographs showing the epidermal structure of the tongue o f: (1) Rat snake (2) Cobra
  • V. Osteology of the skull and the ligaments associated
  • 1. Jaw apparatus
  • 2. Nasal and orbital region
  • 3. Cranium
  • PLATE - VII Skull of tropidonotus.
  • PLATE - Vlll Skull of ratsnake.
  • PLATE - IX Skull of cobra.
  • Quadrato-mandibular joint and the cranio-facial hinge
  • I. Quadrato-mandibular joint
  • Cranio-facial hinge
  • Ligaments of the skull
  • VI. Structure and disposition of the muscles of the feeding apparatus, and its operative efficiency
  • 1.Structure and disposition
  • A. Jaw-muscles
  • 1. Adductors of the lower jaw
  • 1.a. M, Adductor mandibulae externus superficialis
  • PLATE - X Jaw-muscles of tropidonotus.
  • PLATE - XI Jaw-muscles of ratsnake.
  • PLATE - X II Jaw-muscles of cobra.
  • PLATE - XIII Nerve supply to the jaw-muscles of tropidonotus (Fig.1)ratsnake (Fig.2) and cobra (Fig: 3) .
  • PLATE - XIII Nerve supply to the jaw-muscles oftropidonotus (Fig.1)ratsnake (Fig.2) and cobra (Fig: 3) .
  • 1.b. M. Adductor mandibulae externus medialis
  • I. c. Adductor mandibulae externus profundus
  • 1.d. M.Adductor mandibulae posterior
  • 1.e. M.Pseudotemporalis
  • 2. Abductors of the lower jaw
  • 2.a. M.Depressor mandibulae anterior
  • 2.b. M. Depressor mandibulae posterior
  • 3. Protractors of the upper jaw
  • 3.a. M. Protractor pterygoidei
  • 3.b. M. Levator pterygoidei
  • 3.c. M. Protractor quadrati
  • 4. Protractors of the lower jaw and retractors of the upper jaw
  • 4.a. M. Pterygoideus
  • 4.b. M. Pterygoideus accessories
  • 5. Retractors of the upper jaw
  • 5.a. M. Retractor pterygoidei.
  • 5.b. M. Retractor vomeris
  • 5.c. M. Cervico-mandibularis
  • 5.d. M. Neuro-mandibularis
  • B. Hyoid-muscles and Tongue-muscles
  • a. Hyoid-muscles
  • PLATE - XIV Diagrammatic representation of the snake-hyoid muscles and tongue-muscles.
  • b. Tongue-muscles
  • II. Operative efficiency of the feeding apparatus
  • 1. Lever systems of the feeding apparatus
  • 2. Quantitative myology
  • VII. Histochemical observations on the metabolites, some related enzymes and neuromuscular junctions of the major jaw-muscles
  • PLATE - XV T.S.of the jaw-muscles of tropidonotus stained for fat.
  • PLATE - XVI T.S. of the jaw-muscles of ratsnake stained for fat.
  • PLATE - XVII T.S. of the jaw-muscles o f cobra stained for fat.
  • PLATE - XVIII T.S. of the jaw-muscles of tropidonotus stained for glycogen.
  • PLATE - XIX T.S. of the jaw-muscles of ratsnake stained for glycogen.
  • PLATE - XX T.S. of the jaw-muscles of cobra stained for glycogen
  • PLATE -XXI T.S. of the jaw-muscles of tropidonotus stained for SDHactivity.
  • PLATE - XXII T.S. of the jaw-muscles of ratsnake stained for SDH activity
  • PLATE - XXIII T.S. of the jaw-muscles of cobra stained for SDH activity.
  • PLATE - XXIV T.S. of the jaw-muscles of tropidonotus stained for phosphorylase activity.
  • PLATE - XXV T.S. of the jaw-muscles of ratsnke stained for phosphorylase activity.
  • PLATE - XXVI T.S. of the jaw-muscles o f cobra stained for phosphorylase activity.
  • PLATE - XXVII T.S. of the jaw-muscles of tropidonotus showing the distributionof fibres stained for alkali-stabile ATPgase.
  • PLATE - XXVIII T.S. of the jaw-muscles of ratsnake showing the distribution of fibres stained for alkali-stabile ATFase.
  • PLATE - XXIX T.S. of the jaw-muscles of cobra showing the distribution of fibres stained for alkali-stabile ATPase.
  • PLATE - XXX T.S. of the jaw-muscles of tropidonotus showing the distribution of fibres stained for acid-stabile ATPase
  • PLATE - XXXI T.S. of the jaw-muscles of ratsnake showing the distribution of fibres stained for acid-stabile ATPase.
  • PLATE - XXXII T.S. of the jaw-muscles of cobra showing the distribution of fibres stained for acid-stabile ATPase.
  • PLATE - XXXIII T.S. of the jaw-muscles of tropidonotus stained for alkaline phos phosphatase.
  • PLATE - XXXIV T.S. of the jaw-muscles of ratsnake stained for alkaline phosphatase.
  • PLATE - XXXV T.S. of the jaw-muscles of cobra stained for alkaline phosphatase.
  • PLATE - XXXVI T.S. of the jaw-muscles of tropidonotus stained for acid phosphatase.
  • PLATE - XXXVII T.S. of the jaw-muscles of ratsnake stained for acid phosphatase.
  • PLATE - XXXVIII T.S. of the jaw-muscles o f cobra stained for acid phosphatase.
  • PLATE - XXXIX T.S. of the jaw-muscles of tropidonotus showing LDH localization in the mitochondria.
  • PLATE - XL T.S. of the jaw-muscles of ratsnake showing LDH localization in the mitochondria.
  • PLATE - XLI T.S. of the jaw-muscles o f cobra showing LDH localization in the mitochondria..
  • PLATE - XLII T.S. of the jaw-muscles of tropidonotus stained for OC-GPDH.
  • PLATE - XLIII T.S. of the jaw-muscles of ratsnake stained for OC-GPDH.
  • PLATE - XLIV T.S. of the jaw-muscles of cobra stained for OC-GPDH.
  • PLATE - XLV The jaw-muscles of tropidonotus showing the localization of AChE at the neuromuscular junctions.
  • PLATE- XLVl The jaw-muscles of ratsnake showing the localization of AChE at the neuromuscular junctions.
  • PLATE- XLVII The jaw-muscles of cobra showing the localization of AChE at the neuromuscular junctions.
  • PLATE - XLVIII The jaw-muscles of tropidonotus showing the localization of BuChE at the neurontuscular junctions.
  • PLATE - XLIX The jaw-muscles of ratsnake showing the localization of BuChE at the neuromuscular junctions.
  • PLATE - L The jaw-muscles of cobra showing the localization of BuChE at the neurontuscular junctions.
  • ATPase
  • Phosphatases
  • α-Glycerophosphate dehydrogenase
  • Cholinesterases
  • VIII. Discussion
  • 1. Food- getting and feeding behaviour
  • 2. Epidermal structure of the tongue
  • 3. Bony elements and ligaments of the skull
  • 4.. Muscles of the feeding apparatus and its operative efficiency.
  • 1. Muscles of the feeding appartus
  • 2. Operative efficiency
  • 5. General histochemical organization of the jaw-muscles
  • IX. Summary and conclusions
  • X. Works cited